This Practice Note considers the role of depositaries of UK undertakings for collective investment in transferable securities (UK UCITS). It explains the UK regulatory framework, which derives from Directive 2009/65/EC (the UCITS Directive), as amended by Directive 2014/91/EU (UCITS V), and is now set out in UK primary and secondary legislation, assimilated law and the Financial Conduct Authority (FCA) Handbook. It examines the appointment, eligibility, functions, liability and delegation obligations of a UK UCITS depositary. UK legal framework and assimilated law The UCITS Directive and UCITS V were implemented in the UK through a combination of primary legislation in the Financial Services and Markets Act 2000 (FSMA 2000), the UCITS Regulations 2011, SI 2011/1613, and by the FCA throughout its Handbook, but primarily in the Collective Investment Schemes sourcebook (COLL). Commission Delegated Regulation (EU) 2016/438 (UCITS Level 2 Delegated Regulation) has been retained in the UK following the end of the Brexit implementation period and from 1 January 2024 became classified as assimilated law. Assimilated Regulation (EU) 2016/438 applies in the UK subject to the amendments